Systematics of Ameiurus nebulosus (brown bullhead)
        The brown bullhead, Ameiurus nebulosus, is native to much of the eastern Atlantic coast, from Nova Scotia to Florida and as far west as the Mississippi River.  Within its large range, the brown bullhead displays a considerable amount of morphological variation, including colouration, vertebral numbers, and number of pectoral rays.  These differences have lead to a debate over the taxonomic classification of the brown bullhead, which has been informally divided into two subspecies, Ameiurus nebulosus nebulosus in the north and Ameiurus nebulosus marmoratus in the south. The northern form is based on LeSureur's 1819 description of a specimen collected from the Delaware River in Pennsylvania.  In 1955 Hollbrook described another specimen from the Altamaha River in Georgia, which he called Pimelodus marmaratus.  These two descriptions are the basis for the northern and southern and southern subspecies.  The systematic status of these two subspecies has remained problematic to scientists and is in need of revision.  Specimens have been abtained from the U.S. National Museum of Natural History, University of North Carolina-Wilmington, and Auburn University in an attempt to resolve this problem.  It is possible that the extreme morphological differences amongst the northern and southern brown bullheads are due entirely to clinal geographic variation.

Undergraduate students involved with this project:
Erin D. Casey (1999-2000) gave presentation at Virginia Academy of Science Annual Meeting, Radofrd Univ, Radford, VA, May 2000.  Will begin grad school at Eastern Illinois Univ., Dept of Biological Sciences, in Fall 2000.